Online Education Kit: 1995: Physical map of Human Genome Completed

National Human Genome Research Institute

National Institutes of Health
U.S. Department of Health and Human Services


1995: Physical Map of Human Genome Completed

Illustration of STS markerA physical map uses sequence-tagged sites (STSs) as markers to order large segments of DNA. One of the goals of the HGP was to complete a physical map with a marker every 100,000 base pairs by 1998. The map built by 1995 was a significant milestone toward that goal; it contained 15,086 STSs, spaced an average of 199,000 base pairs apart.

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Last Updated: May 9, 2013